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Diptera.info :: Family forums :: Asilidae Forum
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French (Sologne) Tolmerus/Machimus problem. This is Tolmerus calceatus (Meigen 1820)
Quaedfliegh
#1 Print Post
Posted on 05-09-2011 17:37
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Machimus/Tolmerus driving me crazy.

Collected 7 specimen of these creatures. All were caught in Broom (Cytisus sp) on dry soil near Souesmes, Sologne, France. They are similar to T. atricapillus but they are larger, all are about 18-19 mm. Abdomen and thorax dusted golden brown, most hair is yellow, bristles on legs all black, 2-4 bristles on margin scutellum. Beard upper side black, lower side red! Hairs on elongation sternite 8 red! and some black. Postocular bristles red! and black. Legs with lots of red/brown. In appearance and behaviour distinctly different from T. atricapillus. As mentioned they are larger, golden brown and prefer to hang in Broom, more then 50 specimen were hanging there in an area of 50x50 meters, temperature about 30 C. On the bare ground Asilus crabroniformis ruled, over 15 specimen. Keys by Seguy and Engel bring no satisfying solution. Is this maybe a special form?

Edit of tuesday september 6.

Compared my male Loire specimina with 10 randomly picked male specimina of T. atricapillus from the local Museum. I checked them basically for i had little time.

All 10 Dutch specimina had black postocular bristles, a black and white (sometimes yellowish) beard (mystax), and black hairs on elongation of 8th sternite. All but 1 were smaller in size. All Loire males had a black and red beard, more red than black postoculair bristles and red hairs on elongation with some sparse black.
Quaedfliegh attached the following image:


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Edited by Quaedfliegh on 21-03-2015 22:41
Greetings,

Reinoud
 
Quaedfliegh
#2 Print Post
Posted on 06-09-2011 15:08
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Maybe this helps......space between eyes is half of the size of one eye.
Quaedfliegh attached the following image:


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Edited by Quaedfliegh on 08-09-2011 09:23
Greetings,

Reinoud
 
christoophe
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Posted on 07-09-2011 13:07
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Hi Reinoud
Size is not shocking to Tolmerus atricapillus.
I find the picture all the criteria of this species, except the reddish color and facial swelling is low prominent, unless the picture is misleading.
But I do not know what else it could be other that T. atricapillus.
 
Quaedfliegh
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Posted on 07-09-2011 15:24
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Thanks Cristophe! I have to agree but it keeps on bothering me....Size is very variable, i know. But i have seen hundreds of atricapillus in NL. In my collection average size is about 13 mm and none with red hair or bristles. Average of the red haired (7 specimina) is 17mm. All of these share red hair and bristles and golden brown dusting. I also caught a normal one but not in the broom field. It is against the odds. wish there was an expert who had been there to share the experience because i was convinced i collected an entirely other species than T. atricapillus. But when i used keys of Seguy and Engel i ended up there. On the left Redbeard male and on the right Normal male caught in the forest nearby. Please note difference in colour dusting thorax.
Quaedfliegh attached the following image:


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Edited by Quaedfliegh on 10-09-2011 13:27
Greetings,

Reinoud
 
Quaedfliegh
#5 Print Post
Posted on 07-09-2011 15:33
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Following pictures of male and female in their tipical posture and very much alive.
Quaedfliegh attached the following image:


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Edited by Quaedfliegh on 07-09-2011 15:36
Greetings,

Reinoud
 
Quaedfliegh
#6 Print Post
Posted on 07-09-2011 15:35
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Female
Quaedfliegh attached the following image:


[154.23Kb]
Greetings,

Reinoud
 
christoophe
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Posted on 07-09-2011 20:50
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It's not for me Tolmerus atricapillus.
Check if it Machimus chrysitis, first the last sternites and hypopygium.
 
Quaedfliegh
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Posted on 07-09-2011 21:25
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Thanks again Christophe!

Description fits better but there is still a problem. Scutellum marginal bristels: 2-3 with all specimina and black. Hypopygium has not the tipical shape of M. chrysitis...awkward
Greetings,

Reinoud
 
Quaedfliegh
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Posted on 16-09-2011 00:26
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Just to keep it up there Frown Has anybody got a clue?
Greetings,

Reinoud
 
Quaedfliegh
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Posted on 20-09-2011 23:43
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Put them under the microscope again:

Presutural hair: very short, much shorter than average T. atricapillus.

Orbital bristles: Mostly white with T. atricapillus mostly black.
Greetings,

Reinoud
 
Quaedfliegh
#11 Print Post
Posted on 17-10-2011 23:44
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I still want to know! If someone has an idea or question let me know!
Greetings,

Reinoud
 
Quaedfliegh
#12 Print Post
Posted on 30-11-2011 23:15
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I do not know if Tolmerus atricapillus calceatus still exists. Although i'm still convinced this is a different species. Mostly because of different behaviour, posture, size and colour. Smile Is there a Machimus/Tolmerus expert in the house??????
Edited by Quaedfliegh on 30-11-2011 23:16
Greetings,

Reinoud
 
Quaedfliegh
#13 Print Post
Posted on 02-09-2013 21:40
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I think this problem is solved (almost). Update will follow on this solistic thread.
Greetings,

Reinoud
 
Quaedfliegh
#14 Print Post
Posted on 21-03-2015 22:39
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This is Tolmerus calceatus (Meigen, 1820) Publication will follow somewhere in the future Rehabilitation of a species supported by DNA analysis.
Greetings,

Reinoud
 
christoophe
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Posted on 22-03-2015 12:41
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CongratulationsTumbsUp
 
Auratus
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Posted on 23-03-2015 19:57
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Good job Reinoud!
Greetings,

Jan
 
Quaedfliegh
#17 Print Post
Posted on 02-11-2018 17:08
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And here it finally is: https://www.researchgate.net/publication/328642153_Tolmerus_calceatus_Meigen_confirmed_as_a_valid_species_separate_from_Tolmerus_atricapillus_Fallen_Diptera_Asilidae?fbclid=IwAR1hI423wnFma8Lbv-1KQ-b2CsYhhDcNJTfw9gzgSw8iU1-awk1_KeUztU8
Greetings,

Reinoud
 
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