Thread subject: Diptera.info :: Empidoidea from California

Posted by John Carr on 25-09-2013 20:01
#1

These were posted originally to BugGuide and are submitted here with the photographer's permission. They are common on small flowers in the redwood forest of coastal California (USA) early in the year.

I have ruled out everything I know.

At first I thought Oedaleinae (e.g. Anthalia) but the Hybos-like projecting proboscis rules out Ocydromiinae and Oedaleinae. The discal cell emits three veins (see third photo), inconsistent with Hybotinae. Iteaphila and Brachystoma would have R4+5 forked. Empidinae would have a downward projecting proboscis. Clinocerinae has the wrong habitat, forked R4+5, and a weak anal lobe. Hemerodromiinae has modified fore legs. Tachydromiinae lack a discal cell. Anthepiscopus seems to differ in detail, but is at least close. Oreogeton lacks the projecting proboscis.

It is possible that two or more species are represented. The antennae vary.

Four from February 12, 2011:

bugguide.net/images/raw/MQFRIQYRXQFR0Q3RIQTR90DRSQJRP0JRXQARZQURM0YQJK9RZQ9RG0TQFKFQ40JQ50K050NRKQZ0N0.jpg

bugguide.net/images/raw/4QV04QB08QV0EQA0LKB09QLSUQ30GKCK9QZSWQ30WQBK9QB0UQLSUQZS9QCK5KDKIKDKMKRSUQY0AQ.jpg

bugguide.net/images/raw/MLVZ4LBZ8LVZML8Z0L6Z4LLR8L9ZEHLR5HOH8HCH9H3H9H3H4LLR2H3H5L3HWHYHML1ZILFHUH9ZPH.jpg

bugguide.net/images/cache/RL4ZQL5Z8L1H9HCH8HOH4HCH7H5ZSLPZ8LUZRLDHMHTHSLBHXHTHIHCHSLBHHLEZKL1HMHBHMH5Z0LBZWHWZIL1Z7L.jpg
(Click for larger size.)

Two from March 20, 2010:

bugguide.net/images/raw/7HFH2HAHIHTHIHEZIHJH6HYHGL2Z0LVH8HOHGH4ZQLVH7HTHZL6ZGLUZ7LBZ5LJHMH1HXHTH8HDHQL.jpg
bugguide.net/images/raw/3LBLSZ6LYLNLYLULIRKHMZDL0ZBLSZAL0ZZHPRQHMRKHXZHHSRYZ0RDZIR9LKRELKRJZIR6LKRDZHZ.jpg

Later in the year small flowers have similar flies with angled proboscis, probably a different species in Oedaleinae.

bugguide.net/images/cache/OLMZBLXZ2L2RTZXRTZXRJZIROZQRFZJLWLYLNL3L2L0Z2L3LFZHZDZ0RJZRZULXRDZ3L9L3L2LLZ9LRZCZIR9LSR9L.jpg