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That is very interestig, Tony, nice pictures too.

Is tha a general pattern in Ephydridae, that 'aquatic' species have reduced pulvilli?
I had another look at Poecilobothrus: this doly has normal pulvilli and no particularly long hairs on its tarsi.

Though I never really studied it, I've seen all kinds of insects drown...:( so there must be some special adaptations for the water walkers.

Louis