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T. apterygon is c 2mm long. (see Plant & Deeming Int. J. Dipterol. Res 17(1) 13-16 2006). The type locality is a steep mountain slope at 1400m in the Appenine Mountains of Italy at the edge of alpine meadow and deciduous woodland. A single male was found on the ground layer of stones and organic rubbish (how I envy the numbers you are finding in Portugal!)
To my knowledge brachyptery (loss of wings) is only otherwise known in T rossica (described by Igor from Mongolia) while T. brevipennis & T schnitteri have greatly reduced wings. Considering the large number of Tachydromia and their cursorial habits, it is perhaps no surprise that another one has been found. Maybe we should all be searching the ground in mountain areas and might find a few more?
The question of mating habits was raised earlier. I suggect that Tachydromia has more a 'grab and rape' strategy rather than a more complex display component. The related genus Tachypeza sometimes has strange marks on the front legs and these might possibly be used in displaying to females (especially likely as these markings are lacking in the female). However, Tachypeza has more bland fore legs so I still support the grab & rape hypothesis.
Good luck with finding out more about this species:- and if you have any spare specimens.............

cheers
Adrian