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Always interested in single females (!): I think I might make some progress with evaluation of spermathecae. As to the behaviour of males, they are relatively well represented in collections from wine-traps, but rare (as you correctly state) in nature...or our appreciation of nature anyway. It might be that a) they fly higher than our nets reach, b) they don't feed as adults, so don't live very long or c) behave like typical males and hang around on street corners waiting for the girls to go past.