Posted by von Tschirnhaus on 03-09-2024 18:36
#7
Chamaemyiidae, Cremifaniinae, Cremifania nigrocellulata Czerny, 1904. This "aphid fly" was identified by Dieter Doczkal (Munich) in correspondence with me. We both had identified millions of flies during our life, but D.D. found only 1 specimen and I never caught this rarity. It was partly included in an own family Cremifaniidae with only one Nearctic genus and three species. Larvae develop as predators among Adelgidae larvae (compare Zwölfer & Pschorn-Walcher 1960, Zeitschrift für angewandte Entomologie 46: 260-273, not being available in the net). On the first glance, the fly seems to look like a tropical agromyzid with picture wings. But incurved lower orbital bristles are missing (known only in the male of Cerodontha nigricornis in the high Andes, see introduction of https://sdei.senckenberg.de/tschirnhaus-agromyzidae/ ), but orbital setulae inside the upper eye-margin and parafacialia below the eyes (=cheeks) are present. As well, 2 dorsocentrals occur in many agromyzids and vibrissae can be absent in certain leaf-miner flies. Differences are the complete subcosta which ends far basally of the R1 vein. Absent peristomal setae are unknown in agromyzids, and the hind cross-vein stands distinctly more distally compared with agromyzids.