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Thank you very much John. You are correct that South Africa is Zone 4. Zone 1 is an eastern coastal zone that also stretches down into SA. However I think it keys out similarly whichever’s route is taken. What confused me is that G brevipalpis is generally described as being a darker fly without prominent dark spots in the back. However, all else fits it best and some features of G longipennis, the only possible alternative, as this is clearly Fusca group with shorter proboscis (shorter than width of head) just does not fit. I would think general colour is more variable.

See also: Non regional (more complete) key: http://www.fao.org/3/i0535e/i0535e03.pdf

Thanks again.

Best wishes, Philip